Powdermill Nature Reserve
Spring 2003 Banding Summary


Olive-sided Flycatcher, banded at Powdermill May 18, 2003

The bird-banding programat Powdermill Nature Reserve, a 2,000 acre field biological station of Carnegie Museum of Natural History located in the Laurel Highlands region of southwestern Pennsylvania, was in operation on 74 out of a possible 93 days during the spring 2003 season (March 1 through June 1).  A nearly average 2,479 birds of 105 species was banded, with the busiest day being 145 birds of 38 species on May 15--this was the daily maximum for both number of birds and number of species during spring 2003 (Figure 1).   Although not as busy a season as last spring (when 3,475 birds of 109 species were banded), this one day total for number of birds and number of species banded exceeded the highest daily totals recorded last spring (137 birds banded on 4/27/02 and 37 species banded on 5/15/02).   The catch on this spring's busiest day was dominated by wood warblers (16 species), including 27 Magnolia Warblers, 12 Common Yellowthroats, nine American Redstarts, and seven each of Chestnut-sided and Black-throated Blue Warblers.
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Figure 1.  Daily banding totals and species counts for spring 2003 at Powdermill Nature Reserve.  Smaller inset graph shows the relationship between daily banding totals and daily species counts.

Chart Chart2


The timing of spring migrationacross species is what produces the pattern of daily banding totals (both in terms of number birds and number of species) like that shown in Figure 1 above.   Figure 2 below is based on the cumulative spring banding totals for more than 75 species over the last 40 years at Powdermill.  It shows interspecific differences in seasonal occurrence (ranging from the entire spring season, i.e., March 1 through May 31, e.g., for American Robin and American Goldfinch, to little more than two weeks in mid- to late May, e.g.,  for Bay-breasted and Blackpoll warblers), as well as patterns of seasonal abundance for each species (and for the sexes within selected species).  Because the abundance scale for each species is different (and not shown), however, seasonal abundance among species cannot be compared directly.
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The graphs making up Figure 2 depict changes in relative abundance for species across the spring season.  In species for which all or most birds can be sexed, the timing of migration of the sexes is shown separately (males in blue; females in red); species with some or all birds of unknown sex are shown in pale yellow.  In all species, the area of the curves (including for the sexes when they are distinguished) is proportional to abundance for any given banding date.  For example, in Ruby-crowned Kinglet, the first three weeks of the species' migration at Powdermill is entirely comprised of male birds, followed by a week or more when the sexes occur in about equal numbers, and finally by a two week period when virtually all of the RCKIs banded are females.  From Figure 2, is is apparent that, overall, the species is most abundant at Powdermill during the roughly ten-day period in late April when the sexes co-occur in migration.

Figure 2 shows that the early part of the spring migration at the Powdermill banding station (i.e., March through early April) is dominated by captures of winter resident individuals of species like Blue Jay and Black-capped Chickadee and of no more than a dozen or so early migrants, like Golden-crowned Kinglet, Fox and Song sparrows, and Dark-eyed Junco.  The middle portion of the spring season (i.e., the last 2-3 weeks of April) brings the bulk of Ruby-crowned Kinglets and other sparrows, blackbirds, and finches (e.g., Chipping, Field, White-throated, and Swamp sparrows, Brown-headed Cowbird, and Purple Finch), while the height of the migration in mid-May (i.e., 10-20 May; see also Figure 1) coincides with flights of the greatest diversity of migrants, especially Neotropical migrants like Ruby-throated Hummingbirds, vireos, thrushes, orioles, and most wood warblers.  The tail end of the migration (i.e, late May) is dominated by several species of flycatchers and later migrating species of thrushes and wood warblers.  The end of May cutoff for the graphs in Figure 2 clearly cuts off the migration artificially for several of these species, some of which are still migrating through Powdermill as late as mid-June.
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Figure 2.  Timing of spring migration of selected bird species at Powdermill Nature Reserve, based on cumulative daily banding totals from March through May, 1962-2002.








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Totals for all species banded at Powdermill in spring 2003 are given in descending order in
Table 1below.   There were no rarities among the birds banded this spring, but there were some highlights, including our first Olive-sided Flycatcher in several years (photo at top of page).  Some conspicuous "misses" this spring (i.e., species with a long-term average of at least one banded in spring) included Yellow-billed Cuckoo, Warbling Vireo, Golden-winged Warbler, Cape May Warbler, Pine Siskin, and Evening Grosbeak.
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The bird of the season was Magnolia Warbler--this spring's total of 156 MAWAs was our fifth highest ever and more than twice the long-term average (74).  Spring 2003 totals for four species tied or set new record highs: Fox Sparrow (71 banded; ties previous high in 2000); "Traill's" Flycatcher (54 banded; previous record high was 45 in 1996); Chestnut-sided Warbler (35 banded; previous record was 32 in 1990); Northern Parula (7 banded; previous record was 4 in spring 1994).  In contrast, we failed to catch even one Golden-winged Warbler for just the second time in 42 years of spring banding.  Overall, among the 105 species banded this spring, 69 were caught in numbers within one standard deviation of their long-term average, 32 were >1S.D. above average, and just four species were banded in below (<1.S.D.) average numbers.  In general, the spring 2003 flight for Empidonax flycatchers was very strong, with Least, Traill's (i.e., both Willow and Alder, most individuals of which cannot be distinguished in hand), and Yellow-bellied all banded in well above average numbers; many wood warblers also staged good flights, with 16 of 32 species and forms banded in above average numbers and just one, GWWA, in below average numbers.



Table 1.Spring 2003 banding totals by species (in decreasing order of abundance) compared to their long-term average (an "X" denotes if this spring's total is within, greater than, or less than one standard deviation, S.D., of the associated long-term mean).
 
  Comparison with 40-yr. (1962-2002) average (S.D.=standard deviation)
within  1 S.D. >1 S.D. above <1 S.D. below
SPECIES BANDED IN SPRING 2003 
2003
  Totals 
(XX = ties or sets record high or low spring total)
ALL BIRDS 2,479 X    
Magnolia Warbler 156   X  
American Goldfinch 145 X    
Dark-eyed Junco 139     X
Ruby-crowned Kinglet 89 X    
Ruby-throated Hummingbird 87   X  
Nashville Warbler 82   X  
Cedar Waxwing 81 X    
White-throated Sparrow 79 X    
Song Sparrow 73     X
Fox Sparrow 71   XX  
Chipping Sparrow 69 X    
Gray Catbird 68 X    
Common Yellowthroat 67 X    
Least Flycatcher 59   X  
Swamp Sparrow 59   X  
Traill's (Willow and Alder) Flycatcher 54   XX  
American Redstart 52   X  
Indigo Bunting 51   X  
Canada Warbler 42 X    
Yellow Warbler 41 X    
Red-eyed Vireo 39 X    
Blue Jay 39 X    
Red-winged Blackbird 39 X    
Chestnut-sided Warbler 35   XX  
Northern Waterthrush 34   X  
Northern Cardinal 31 X    
House Finch 29 X    
Wilson's Warbler 28 X    
Swainson's Thrush 27 X    
Golden-crowned Kinglet 26   X  
Philadelphia Vireo 25   X  
Field Sparrow 25 X    
Yellow-bellied Flycatcher 24   X  
Eastern Towhee 21 X    
Barn Swallow 20 X    
White-crowned Sparrow 20   X  
Louisiana Waterthrush 19 X    
Mourning Warbler 19 X    
Wood Thrush 18 X    
Hooded Warbler 18 X    
Rose-breasted Grosbeak 17 X    
Tennessee Warbler 16 X    
Tree Swallow 15   X  
American Robin 15 X    
Brown-headed Cowbird 15 X    
Black-and-white Warbler 14   X  
Kentucky Warbler 14   X  
Baltimore Oriole 14   X  
Black-throated Blue Warbler 13   X  
Blue-winged Warbler 12   X  
Yellow-rumped Warbler 12 X    
Lincoln's Sparrow 12 X    
Northern Flicker 11 X    
Black-capped Chickadee 11 X    
Blue-gray Gnatcatcher 11 X    
Mourning Dove 9 X    
Blackpoll Warbler 9   X  
Rusty Blackbird 9 X    
White-eyed Vireo 8 X    
Brown Thrasher 7 X    
Northern Parula 7   XX  
Black-throated Green Warbler 7   X  
Ovenbird 6 X    
Scarlet Tanager 6 X    
Spotted Sandpiper 5 X    
Acadian Flycatcher 5 X    
Eastern Phoebe 5 X    
Northern Rough-winged Swallow 5 X    
Tufted Titmouse 5 X    
Winter Wren 5   X  
Bay-breasted Warbler 5 X    
Purple Finch 5 X    
Solitary Sandpiper 4 X    
Gray-cheeked Thrush 4 X    
Black-billed Cuckoo 3 X    
Belted Kingfisher 3   X  
Eastern Wood-Pewee 3 X    
Blue-headed Vireo 3   X  
Eastern Bluebird 3 X    
Hermit Thrush 3   X  
Worm-eating Warbler 3   X  
American Tree Sparrow 3 X    
House Sparrow 3 X    
Red-bellied Woodpecker 2 X    
Yellow-bellied Sapsucker 2 X    
Brown Creeper 2 X    
House Wren 2     X
Orange-crowned Warbler 2 X    
Blackburnian Warbler 2   X  
Yellow-throated Warbler 2   X  
Prairie Warbler 2 X    
"Yellow" Palm Warbler 2   X  
Yellow-breasted Chat 2 X    
Common Grackle 2 X    
Orchard Oriole 2 X    
American Woodcock 1 X    
Northern Saw-whet Owl 1 X    
Downy Woodpecker 1     X
Hairy Woodpecker 1 X    
Olive-sided Flycatcher 1 X    
White-breasted Nuthatch 1 X    
Veery 1 X    
European Starling 1 X    
"Western" Palm Warbler 1 X    
Grasshopper Sparrow 1 X    
SPECIES MISSED* IN SPRING 2003  
Yellow-billed Cuckoo 0 X    
Warbling Vireo 0 X    
Golden-winged Warbler 0     XX
Cape May Warbler 0 X    
Pine Siskin 0 X    
Evening Grosbeak 0 X    
* Species with an average spring banding total >1


The real highlight of the spring 2003 banding season wasn't a bird but an event, the 80th annual meeting of the Eastern Bird Banding Association, held at Powdermill and nearby venues from 4-6 April.  This meeting, attended by nearly 100 people, was followed by a sanctioned North American Banding Council bander level certification workshop from 6-8 April.  Both events were a great success despite the fact that the prevailing weather on both occasions was mostly inclement!  A very special guest at the EBBA meeting was Dr. Lukas Jenni, Scientific Director of the Swiss Ornithological Institute and co-author of the extraordinary Moult and Ageing of European Passerines, who was the meeting's keynote speaker and who also gave a very informative and  interesting workshop on the European ringing (=banding) schemes and techniques.  Another special guest, British ringer, Dr. David Norman, contributed much of significance to both the workshops and the scientific paper sessions at the EBBA meeting, and he stayed on to provide invaluable assistance during the NABC certification session.   All of us here at Powdermill, and all those in attendance at these events, were signally fortunate in having the opportunity to "expand our worlds of banding" by meeting and learning from these extraordinarily knowledgeable gentlemen.  We can't thank them enough for their participation--they truly made these events a once-in-a-lifetime experience for all of us!



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We conclude this banding summary with our sincere thanks to the following volunteers, interns, and others who helped us greatly with the banding this spring:  Randi Gerrish, Jim Gruber, Brian Jones, Carroll Labarthe, Mike Lanzone, Adrienne Leppold, David Liebmann, Mike Lutmerding, Annie LindsayMarilyn Niedermeier, David Norman, Diane Pick, Katharine Reinemann, Carole Shanahan, Joe Schreiber, and Nathan Tarr.  Our special thanks to Brad Silfies for making arrangements to have this year's EBBA meeting at Powdermill and to fellow NABC Trainers Betsy Brooks, Mary Doscher, Mary Gustafson, Sara Morris, and Hannah Suthers, whose individual and collective organizational and other skills the Powdermill banders relied on so heavily during the NABC certification session!


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